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Copyeditor: Jo Clayton Proofreader: Sally Huish Indexer: Bill Johncocks Permissions Coordinator: Sheri Gilbert Back Cover Photograph: Photography, Christophe Carlinet; Design, Nigel Orme Molecular Biology of the Cell Interactive Media: Artistic and Scientific Direction: Peter Walter Narration: Julie Theriot Director of Digital Publishing: Michael Morales Editorial Assistant: Leah Christians Production Editor: Natasha Wolfe 2015 by Bruce Alberts, Alexander Johnson, Julian Lewis, David Morgan, Martin Raff, Keith Roberts, and Peter Walter.

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About the Authors Bruce Alberts received his PhD from Harvard University and is the Chancellors Leadership Chair in Biochemistry and Biophysics for Science and Education, University of California, San Francisco.

He was the editor-in-chief of Science magazine from 2008 until 2013, and for twelve years he served as President of the U.S.

National Academy of Sciences (1993 2005).

Alexander Johnson received his PhD from Harvard University and is Professor of Microbiology and Immunology at the University of California, San Francisco.

Julian Lewis (19462014) received his DPhil from the University of Oxford and was an Emeritus Scientist at the London Research Institute of Cancer Research UK.

David Morgan received his PhD from the University of California, San Francisco, and is Professor of the Department of Physiology there as well as the Director of the Biochemistry, Cell Biology, Genetics, and Developmental Biology Graduate Program.

Martin Raff received his MD from McGill University and is Emeritus Professor of Biology at the Medical Research Council Laboratory for Molecular Cell Biology at University College London.

Keith Roberts received his PhD from the University of Cambridge and was Deputy Director of the John Innes Centre, Norwich.

Peter Walter received his PhD from the Rockefeller University in New York and is Professor of the Department of Biochemistry and Biophysics at the University of California, San Francisco, and an Investigator at the Howard Hughes Medical Institute.

John Wilson received his PhD from the California Institute of Technology and pursued his postdoctoral work at Stanford University.

He is Distinguished Service Professor of Biochemistry and Molecular Biology at Baylor College of Medicine in Houston.

Tim Hunt received his PhD from the University of Cambridge where he taught biochemistry and cell biology for more than 20 years.

He worked at Cancer Research UK until his retirement in 2010.

He shared the 2001 Nobel Prize in Physiology or Medicine with Lee Hartwell and Paul Nurse.